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fumarate 4-maleylacetoacetate 4-hydroxyphenylpyruvate homogentisate 4-fumarylacetoacetate O2 L-tyrosine H2O O2 L-glutamate acetoacetate CO2 2-oxoglutarate 1.13.11.5 | homogentisate 1,2-dioxygenase 2.6.1.5 | tyrosine transaminase 5.2.1.2 | maleylacetoacetate isomerase 1.13.11.27 | 4-hydroxyphenylpyruvate dioxygenase 3.7.1.2 | fumarylacetoacetase